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    Catasetum , a genus discussed briefly by Darwin , actually launches its viscid pollinia with explosive force when an insect touches a seta , knocking the pollinator off the flower.

    After pollination, the sepals and petals fade and wilt, but they usually remain attached to the ovary. Some species, such as Phalaenopsis , Dendrobium , and Vanda , produce offshoots or plantlets formed from one of the nodes along the stem , through the accumulation of growth hormones at that point.

    These shoots are known as keiki. The ovary typically develops into a capsule that is dehiscent by three or six longitudinal slits, while remaining closed at both ends.

    The seeds are generally almost microscopic and very numerous, in some species over a million per capsule. After ripening, they blow off like dust particles or spores.

    They lack endosperm and must enter symbiotic relationships with various mycorrhizal basidiomyceteous fungi that provide them the necessary nutrients to germinate, so all orchid species are mycoheterotrophic during germination and reliant upon fungi to complete their lifecycles.

    As the chance for a seed to meet a suitable fungus is very small, only a minute fraction of all the seeds released grow into adult plants. In cultivation, germination typically takes weeks.

    Horticultural techniques have been devised for germinating orchid seeds on an artificial nutrient medium, eliminating the requirement of the fungus for germination and greatly aiding the propagation of ornamental orchids.

    The usual medium for the sowing of orchids in artificial conditions is agar gel combined with a carbohydrate energy source.

    The carbohydrate source can be combinations of discrete sugars or can be derived from other sources such as banana , pineapple , peach , or even tomato puree or coconut water.

    After the preparation of the agar medium, it is poured into test tubes or jars which are then autoclaved or cooked in a pressure cooker to sterilize the medium.

    After cooking, the medium begins to gel as it cools. The taxonomy of this family is in constant flux, as new studies continue to clarify the relationships between species and groups of species, allowing more taxa at several ranks to be recognized.

    Five subfamilies are recognised. The cladogram below was made according to the APG system of It represents the view that most botanists had held up to that time.

    It was supported by morphological studies , but never received strong support in molecular phylogenetic studies. Apostasioideae : 2 genera and 16 species, south-western Asia.

    Cypripedioideae : 5 genera and species, from the temperate regions of the world, as well as tropical America and tropical Asia. Vanilloideae : 15 genera and species, humid tropical and subtropical regions, eastern North America.

    Epidendroideae : more than genera and more or less 20, species, cosmopolitan. Orchidoideae : genera and 3, species, cosmopolitan. In , a phylogenetic study [12] showed strong statistical support for the following topology of the orchid tree , using 9 kb of plastid and nuclear DNA from 7 genes , a topology that was confirmed by a phylogenomic study in the same year.

    A study in the scientific journal Nature has hypothesised that the origin of orchids goes back much longer than originally expected.

    The bee was carrying pollen of a previously unknown orchid taxon, Meliorchis caribea , on its wings. This find is the first evidence of fossilised orchids to date [14] and shows insects were active pollinators of orchids then.

    This extinct orchid, M. Genetic sequencing indicates orchids may have arisen earlier, 76 to 84 million years ago during the Late Cretaceous.

    Chase et al. Using the molecular clock method, it was possible to determine the age of the major branches of the orchid family. This also confirmed that the subfamily Vanilloideae is a branch at the basal dichotomy of the monandrous orchids, and must have evolved very early in the evolution of the family.

    Since this subfamily occurs worldwide in tropical and subtropical regions, from tropical America to tropical Asia, New Guinea and West Africa, and the continents began to split about million years ago, significant biotic exchange must have occurred after this split since the age of Vanilla is estimated at 60 to 70 million years.

    Genome duplication occurred prior to the divergence of this taxon. There are around genera of orchids. The following are amongst the most notable genera of the orchid family: [ citation needed ].

    The type genus i. In Middle English , the name bollockwort was used for some orchids, based on " bollock " meaning testicle and " wort " meaning plant.

    Orchidaceae are cosmopolitan , occurring in almost every habitat apart from glaciers. The following list gives a rough overview of their distribution: [ citation needed ].

    A majority of orchids are perennial epiphytes , which grow anchored to trees or shrubs in the tropics and subtropics.

    Species such as Angraecum sororium are lithophytes , [24] growing on rocks or very rocky soil. Other orchids including the majority of temperate Orchidaceae are terrestrial and can be found in habitat areas such as grasslands or forest.

    Some orchids, such as Neottia and Corallorhiza , lack chlorophyll , so are unable to photosynthesise. Instead, these species obtain energy and nutrients by parasitising soil fungi through the formation of orchid mycorrhizae.

    The fungi involved include those that form ectomycorrhizas with trees and other woody plants, parasites such as Armillaria , and saprotrophs.

    While only a few species are achlorophyllous holoparasites , all orchids are myco-heterotrophic during germination and seedling growth, and even photosynthetic adult plants may continue to obtain carbon from their mycorrhizal fungi.

    The other important use of orchids is their cultivation for the enjoyment of the flowers. Most cultivated orchids are tropical or subtropical , but quite a few that grow in colder climates can be found on the market.

    Temperate species available at nurseries include Ophrys apifera bee orchid , Gymnadenia conopsea fragrant orchid , Anacamptis pyramidalis pyramidal orchid and Dactylorhiza fuchsii common spotted orchid.

    Orchids of all types have also often been sought by collectors of both species and hybrids. Many hundreds of societies and clubs worldwide have been established.

    These can be small, local clubs, or larger, national organisations such as the American Orchid Society. Both serve to encourage cultivation and collection of orchids, but some go further by concentrating on conservation or research.

    The term "botanical orchid" loosely denotes those small-flowered, tropical orchids belonging to several genera that do not fit into the "florist" orchid category.

    A few of these genera contain enormous numbers of species. Some, such as Pleurothallis and Bulbophyllum , contain approximately and species, respectively, and are often extremely vegetatively diverse.

    The primary use of the term is among orchid hobbyists wishing to describe unusual species they grow, though it is also used to distinguish naturally occurring orchid species from horticulturally created hybrids.

    The dried seed pods of one orchid genus, Vanilla especially Vanilla planifolia , are commercially important as a flavouring in baking , for perfume manufacture and aromatherapy.

    The underground tubers of terrestrial orchids [mainly Orchis mascula early purple orchid ] are ground to a powder and used for cooking, such as in the hot beverage salep or in the Turkish frozen treat dondurma.

    The similarity in appearance to testes naturally accounts for salep being considered an aphrodisiac. The dried leaves of Jumellea fragrans are used to flavour rum on Reunion Island.

    Some saprophytic orchid species of the group Gastrodia produce potato-like tubers and were consumed as food by native peoples in Australia and can be successfully cultivated, notably Gastrodia sesamoides.

    Wild stands of these plants can still be found in the same areas as early Aboriginal settlements, such as Ku-ring-gai Chase National Park in Australia.

    Aboriginal peoples located the plants in habitat by observing where bandicoots had scratched in search of the tubers after detecting the plants underground by scent.

    Orchids have many associations with symbolic values. For example, the orchid is the City Flower of Shaoxing , China. Cattleya mossiae is the national Venezuelan flower, while Cattleya trianae is the national flower of Colombia.

    Vanda 'Miss Joaquim' is the national flower of Singapore , Guarianthe skinneri is the national flower of Costa Rica and Rhyncholaelia digbyana is the national flower of Honduras.

    Panama 's national flower is the Holy Ghost orchid Peristeria elata , or 'the flor del Espiritu Santo'. Rhynchostylis retusa is the state flower of the Indian state of Assam where it is known as Kopou Phul.

    Orchids native to the Mediterranean are depicted on the Ara Pacis in Rome, until now the only known instance of orchids in ancient art, and the earliest in European art.

    Cattleya mossiae ' Empress Frederick'. From Wikipedia, the free encyclopedia. The orchid family of flowering plants.

    For other uses, see Orchid disambiguation. Main article: Taxonomy of the Orchid family. Main article: List of natural Orchidaceae genera.

    Further information: Vanilla. Cattleya Queen Sirikit 'Diamond Crown'. Botanical Journal of the Linnean Society. Archived from the original PDF on 25 May Retrieved 26 June World Checklist of Selected Plant Families.

    Retrieved 2 April See External links below. Chase Conservation Biology. Complete guide to orchids. Meredith Publishing Group p.

    Orchids in Northern Washington State. Silvercrown Mountain Outdoor School. Retrieved 10 June Australian Geographic.

    Archived from the original on 30 November Retrieved 26 May Journal of Chemical Ecology. Animal Behaviour. Behavioral Ecology and Sociobiology.

    Renner New Phytologist. Mark; Williams, Norris H. Proceedings of the Royal Society, series B. Singer; Charles R.

    Marshall; Naomi E. Pierce 30 August Rasmussen Pridgeon; P. Cribb; M. Chase; F. Rasmussen eds. Orchidoideae Part 1. Genera Orchidacearum.

    Oxford University Press. Oxford: Oxford University Press. A Dictionary of English Plant Names. London: Allen Lane. Oxford English Dictionary.

    Retrieved 19 January Leake

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